Information

Plants terrestrial or epilithic. Rhizome short-decumbent, to 30 mm long, to 5 mm in diameter, set with roots, closely spaced persistent stipe bases and scales, scales thinly crustaceous, ferrugineus, adnate, linear to subulate, irregularly and shallowly dentate, apex terminates in a small thin-walled cell or a short subulate cell, to 6 mm long, to 0.5 mm wide. Fronds crowded, erect, to 435 mm long, to 7 per plant; stipe rigid, atrocastaneus, terete, to 150 mm long, to 2 mm in diameter, initially densely scaled, scales chartaceus, stramineous to ferrugineus, sessile, filiform to subulate, entire, apex terminates in a small thin-walled cell or a short subulate cell, to 5 mm long, to 0.3 mm wide; lamina oblong-acute, 1-pinnate, to 300 mm long, to 90 mm wide, with up to 52 pinna pairs; rachis atrocastaneus, terete, initially closely set with hairs and scales similar to those on the stipe; pinnae petiolate, petiole to 2 mm long, closely set with hairs and scales similar to those on rachis, opposite to alternate, pectinate, generally more widely spaced basally, coriaceus, linear, entire, cordate to inaequilaterally cordate, acute, to 44 mm long, to 3 mm wide, glabrous adaxially, abaxially with filiform scales and pluricellular, uniseriate hairs along costa, to 2 mm long, articulated; costa adaxially sulcate. Venation obscure, forked, free, branches terminate near the margin. Sori forms a continuous line along a marginal vascular commissure; receptacle with simple, pluricellular, uniseriate hairs terminating in an orange oblong to clavate cell, to 0.5 mm long; sporangium long-stalked, stalk simple, 3-seriate below capsule, capsule globose to obovate in lateral view, with (14-)14(-16) indurated annulus cells, epistomium (4-)4(-6)-celled, hypostomium (4-)4(-6)-celled, stomium with 4 narrow cells with conspicuously thickened walls; indusium continuous, near-marginal, chartaceus, entire, to 0.4 mm wide. Spores 32 per sporangium, brown, smooth-granulose, monolete, elliptic, (66-)68(-72) x (44-)45.33(-48) (m, or tetrahedral-globose, trilete, (50-)54(-64) (m. Figure 19E & F.