Plants terrestrial or epilithic. Rhizome short-decumbent, closely branched, to 2 mm in diameter, closely set with hairs, hairs crustaceous, atrocastaneus, basally attached, pluricellular, uniseriate, to 1 mm long. Fronds caespitose, erect, closely spaced, to 3 mm apart, to 350 mm long; stipe black, terete, narrowly winged along the entire length, to 185 mm long, to 1.2 mm in diameter, sparsely haired, hairs crustaceous, ventrally attached, subulate, 2 to 3-celled, to 0.3 mm long; lamina anadromous, to 3-pinnate, lanceolate, ovate, or narrowly elliptic, to 165 mm long, to 85 mm wide, with up to 17 pinna pairs; rachis terete, narrowly winged, sparsely set with hairs similar to those on the stipe; pinnae petiolate, petiole to 1 mm long, thinly herbaceous, opposite to alternate, usually slightly overlapping, to 2-pinnate, lanceolate to narrowly ovate, to 57 mm long, to 17 mm wide, with up to 9 pinnule pairs; pinna-rachis narrowly winged, sparsely set with hairs similar to those on the stipe; pinnules alternate, sessile, spaced or slightly overlapping, 1-pinnate, lanceolate, narrowly trullate, or oblong, to 15 mm long, to 5 mm wide; segments narrowly rhomboid to oblong, to 4 mm long, to 2 mm wide, lobed, lobes to 0.3 mm wide, acute, entire. Venation anadromous, evident, ending near the margin. Sori borne on acroscopic anadromous vein branches of the pinnules; involucres turbinate, mouth slightly dilated, to 1.5 mm deep, lobes truncate to obtuse; receptacle protrudes far, to 4 mm long. Sporangium sessile, annulus oblique, complete, with (16-)18(-20) indurated annulus cells, stomium 2- to 5-celled. Spores green, spheroidal, trilete, 33 x 43 µm. Figure 12L & M.