Information

Plants terrestrial or epilithic. Rhizome short-creeping, to 6 mm in diameter, set with roots, closely set persistent stipe bases, and scales, scales concolorous or bicolorous, concolorous scales, chartaceus, stramineous to ferrugineus, bicolorous scales with a crustaceous, castaneus, multistratose central region, margins stramineous, sessile, subulate to subulate-caudate, entire, apex terminates in an oblong thin-walled cell, to 5 mm long, to 1 mm wide. Fronds closely spaced, to 8 per plant, erect, to 540 mm long; stipe castaneus, proximally terete, sulcate in upper half, sulcus flat-bottomed, to 295 mm long, to 3 mm in diameter, proximally with scales similar to those on the rhizome, higher up sparsely set with stramineous, filiform scales and hairs, apex terminates in an oblong thin-walled cell, to 5 mm long, soon becoming glabrous; lamina anadromous, firmly herbaceous to thinly coriaceus, pentagonal, to 3-pinnate-pinnatifid, to 230 mm long, to 170 mm wide, with up to 12 petiolated pinna pairs; rachis and lower order axes castaneus, sulcate, sulci confluent, glabrous; pinnae petiolate, petiole to 14 mm long, opposite to alternate, widely spaced proximally, more closely spaced distally, overlapping, to 2-pinnate-pinnatifid, basiscopically developed, inaequilaterally broadly ovate to ovate, to 130 mm long, to 88 mm wide, with up to 10 petiolated pinnule pairs; pinnules petiolate, petiole to 5.5 mm long, alternate, spaced to overlapping slightly, to 1-pinnate-pinnatifid, basiscopically developed, inaequilaterally broadly ovate to ovate, acroscopic pinnule on basal pinnae to 25 mm long, basiscopic pinnule on basal pinnae to 50 mm long, with up to 5 pairs of petiolated segments; segments opposite to alternate, spaced, short-stalked, pinnatifid, narrowly ovate to oblong-obtuse, to 14 mm long, to 7 m wide; ultimate segments sessile, oblong-obtuse, shallowly lobed, to 3 mm long, to 2 mm wide, glabrous adaxially, abaxially closely set with clavate, 2-celled hairs, to 0.15 mm long. Venation obscure, pinnately branched, free vein branches end in the teeth near the margin. Sori at free vein endings of ultimate segments, marginal, discrete; receptacle nude; sporangium short-stalked, simple, 3-seriate below capsule, capsule circular in lateral view, with (14-)15(-16) indurated annulus cells, epistomium (4-)5(-7)-celled, hypostomium (4-)5(-6)-celled, stomium composed of 5-6 narrow cells of which walls are more thickened than those of surrounding cells; indusium membranous, pale, semi-circular, entire to lacerate. Spores 64 per sporangium, yellowish-brown, tetrahedral-globose, trilete, smoothly rugulose, exospore (34-)38.7(-42) (m in equatorial diameter. Figure 17A-C.

Information

Plants terrestrial or epilithic. Rhizome short-creeping, to 6 mm in diameter, set with roots, closely set persistent stipe bases, and scales, scales concolorous or bicolorous, concolorous scales, chartaceus, stramineous to ferrugineus, bicolorous scales with a crustaceous, castaneus, multistratose central region, margins stramineous, sessile, subulate to subulate-caudate, entire, apex terminates in an oblong thin-walled cell, to 5 mm long, to 1 mm wide. Fronds closely spaced, to 8 per plant, erect, to 540 mm long; stipe castaneus, proximally terete, sulcate in upper half, sulcus flat-bottomed, to 295 mm long, to 3 mm in diameter, proximally with scales similar to those on the rhizome, higher up sparsely set with stramineous, filiform scales and hairs, apex terminates in an oblong thin-walled cell, to 5 mm long, soon becoming glabrous; lamina anadromous, firmly herbaceous to thinly coriaceus, pentagonal, to 3-pinnate-pinnatifid, to 230 mm long, to 170 mm wide, with up to 12 petiolated pinna pairs; rachis and lower order axes castaneus, sulcate, sulci confluent, glabrous; pinnae petiolate, petiole to 14 mm long, opposite to alternate, widely spaced proximally, more closely spaced distally, overlapping, to 2-pinnate-pinnatifid, basiscopically developed, inaequilaterally broadly ovate to ovate, to 130 mm long, to 88 mm wide, with up to 10 petiolated pinnule pairs; pinnules petiolate, petiole to 5.5 mm long, alternate, spaced to overlapping slightly, to 1-pinnate-pinnatifid, basiscopically developed, inaequilaterally broadly ovate to ovate, acroscopic pinnule on basal pinnae to 25 mm long, basiscopic pinnule on basal pinnae to 50 mm long, with up to 5 pairs of petiolated segments; segments opposite to alternate, spaced, short-stalked, pinnatifid, narrowly ovate to oblong-obtuse, to 14 mm long, to 7 m wide; ultimate segments sessile, oblong-obtuse, shallowly lobed, to 3 mm long, to 2 mm wide, glabrous adaxially, abaxially closely set with clavate, 2-celled hairs, to 0.15 mm long. Venation obscure, pinnately branched, free vein branches end in the teeth near the margin. Sori at free vein endings of ultimate segments, marginal, discrete; receptacle nude; sporangium short-stalked, simple, 3-seriate below capsule, capsule circular in lateral view, with (14-)15(-16) indurated annulus cells, epistomium (4-)5(-7)-celled, hypostomium (4-)5(-6)-celled, stomium composed of 5-6 narrow cells of which walls are more thickened than those of surrounding cells; indusium membranous, pale, semi-circular, entire to lacerate. Spores 64 per sporangium, yellowish-brown, tetrahedral-globose, trilete, smoothly rugulose, exospore (34-)38.7(-42) (m in equatorial diameter. Figure 17A-C.