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Plants terrestrial, epilithic or epiphytic. Rhizome wide-creeping, irregularly laterally branched, to 3 mm in diameter, set with roots, widely spaced persistent phyllopodia to 20 mm long, and scales, scales chartaceus, castaneus to ferrugineous, peltate, circular to subcircular, margins regularly set with uni- and bicellular capitate hairs, adaxial surface closely set with filiform uni- or pluricellular hairs, to 2 mm long, to 1.5 mm wide. Fronds erect to arching, widely spaced, articulated to phyllopodia, to 60 mm apart, to 765 mm long; stipe articulated to the phyllopodia, firm, proximally castaneus, stramineous higher up, adaxially sulcate, to 235 mm long, to 2.5 mm in diameter, set with hairs and scales, hairs pluricellular, acicular, or 2 to 3-celled and clavate, to 120 (m long, sparsely scaled, scales chartaceus, ferrugineous, adnate to sessile, broadly ovate, cordate, margins regularly set with unicellular, thin-walled cells, to 2 mm long, to 1.8 mm wide; lamina catadromous, 1-pinnate-pinnatifid, narrow-elliptic, to 520 mm long, to 175 mm wide, with up to 22 sessile pinna pairs; rachis adaxially sulcate, closely set with hairs similar to those on the stipe; pinnae opposite to alternate, more widely spaced and often slightly reduced basally, oblong-cuneate, pinnatifid, adnate and basiscopically decurrent towards the apex, to 85 mm long, to 27 mm wide; costa adaxially convex, densely set with hairs similar to those on the rachis; pinnules herbaceous, adnate, oblong-acute to oblong-obtuse, basiscopically decurrent, 4 mm long, to 6.5 mm in diameter, dentate, adaxially and abaxially regularly set with acicular hairs to 1 mm long along the veins and margin, and with 3-celled clavate hairs on the lamina between the veins, the 2 apical cells being shorter and gland-like, to 110 (m long. Venation evident, pinnately branched, vein branches terminate in a hydathode in the teeth near the margin, often exuding a white crystalline substance adaxially. Sori circular, to 1.2 mm in diameter, terminally on an abbreviated anadromous vein branch; sporangium long-stalked, simple, 3-seriate below the capsule, capsule globose in lateral view, with (10-)13(-15) indurated annulus cells, epistomium (3-)4(-4)-celled, hypostomium 4-celled. Indusium chartaceus, brown, reniform, margin and surface closely set with unicellular capitate gland-like hairs, to 1 mm in diameter. Spores 64 per sporangium, elliptic to spheroidal, monolete, brown, with narrow reticulate wings, exospore (36-)38.08(-42) x (26-)27.13(-32) (m. Figure 42A & B.

Information

Plants terrestrial, epilithic or epiphytic. Rhizome wide-creeping, irregularly laterally branched, to 3 mm in diameter, set with roots, widely spaced persistent phyllopodia to 20 mm long, and scales, scales chartaceus, castaneus to ferrugineous, peltate, circular to subcircular, margins regularly set with uni- and bicellular capitate hairs, adaxial surface closely set with filiform uni- or pluricellular hairs, to 2 mm long, to 1.5 mm wide. Fronds erect to arching, widely spaced, articulated to phyllopodia, to 60 mm apart, to 765 mm long; stipe articulated to the phyllopodia, firm, proximally castaneus, stramineous higher up, adaxially sulcate, to 235 mm long, to 2.5 mm in diameter, set with hairs and scales, hairs pluricellular, acicular, or 2 to 3-celled and clavate, to 120 (m long, sparsely scaled, scales chartaceus, ferrugineous, adnate to sessile, broadly ovate, cordate, margins regularly set with unicellular, thin-walled cells, to 2 mm long, to 1.8 mm wide; lamina catadromous, 1-pinnate-pinnatifid, narrow-elliptic, to 520 mm long, to 175 mm wide, with up to 22 sessile pinna pairs; rachis adaxially sulcate, closely set with hairs similar to those on the stipe; pinnae opposite to alternate, more widely spaced and often slightly reduced basally, oblong-cuneate, pinnatifid, adnate and basiscopically decurrent towards the apex, to 85 mm long, to 27 mm wide; costa adaxially convex, densely set with hairs similar to those on the rachis; pinnules herbaceous, adnate, oblong-acute to oblong-obtuse, basiscopically decurrent, 4 mm long, to 6.5 mm in diameter, dentate, adaxially and abaxially regularly set with acicular hairs to 1 mm long along the veins and margin, and with 3-celled clavate hairs on the lamina between the veins, the 2 apical cells being shorter and gland-like, to 110 (m long. Venation evident, pinnately branched, vein branches terminate in a hydathode in the teeth near the margin, often exuding a white crystalline substance adaxially. Sori circular, to 1.2 mm in diameter, terminally on an abbreviated anadromous vein branch; sporangium long-stalked, simple, 3-seriate below the capsule, capsule globose in lateral view, with (10-)13(-15) indurated annulus cells, epistomium (3-)4(-4)-celled, hypostomium 4-celled. Indusium chartaceus, brown, reniform, margin and surface closely set with unicellular capitate gland-like hairs, to 1 mm in diameter. Spores 64 per sporangium, elliptic to spheroidal, monolete, brown, with narrow reticulate wings, exospore (36-)38.08(-42) x (26-)27.13(-32) (m. Figure 42A & B.