Edit History
Asplenium inaequilaterale Willd. [family ASPLENIACEAE]
Date Updated: 19 August 2007
Herbarium
South African National Biodiversity Institute, Compton Herbarium, Cape Town (SAM)
Collection
Swaziland ferns and fern allies
Resource Type
Reference Sources
Entry From
Swaziland Ferns and Fern Allies, (2003) Author: J.P. Roux
Names
Asplenium inaequilaterale Willd. [family ASPLENIACEAE]
Common names
Asplenium brachyotus Kunze in Linnaea 10: 512 (1836). Asplenium erectum Bory ex Willd. var. brachyotus (Kunze) Sim, Ferns S. Afr.: 138, LXVI, t. 1 (1892). Asplenium laetum Sw. var. brachyotus (Kunze) Bonap., Notes ptérid.16: 60 (1925). Type: Ad catarractum magnam in praeruptis umbrosis inter Omsamcaba, 200 m, Drège s.n. [LZ†, B!, lecto., designated by Roux (1986); BM!, K!, isolecto.]. Asplenium laetum sensu Sim, Ferns S. Afr., 2nd edn: 150, pl. 50. t. 1 (1915), non Sw. (1806).
inaequilateralis = with unequal sides
Vernacular names: Lolly spleenwort; Wortelknop tralievaring (Afr.)
Information
Plants epilithic. Rhizome short, erect to suberect, to 20 mm long, to 4 mm in diameter, set with roots, crowded persistent stipe bases and scales, scales crustaceous, atrocastaneus to black, clathrate, sessile, subulate to lanceolate, cordate to cordate-imbricate, entire or with scattered, pluricellular recurved outgrowths along the margin, apex terminates in a thin-walled cell, to 5 mm long, to 0.7 mm wide. Fronds crowded, caespitose, erect, to 430 mm long; stipe firm, castaneus, matt, terete, to 210 mm long, to 1.6 mm in diameter, narrowly green-winged to the base, base sparsely set with scales similar to those on the rhizome, higher up with filiform clathrate scales terminating in an elliptic thin-walled cell, to 1 mm long, and 3-4-celled hairs terminating in an enlarged obovate thin-walled cell, to 0.3 mm long; lamina 1-pinnate, narrowly elliptic, to 230 mm long, to 68 mm wide, with up to 17 petiolated pinna pairs; rachis firm, castaneus to greenish higher up, shallowly sulcate adaxially, narrowly green-winged, set with filiform scales and hairs similar to those on the stipe, glabrous later; pinnae basal pinnae petiolate, petiole to 1.5 mm long, sessile towards the apex, herbaceous, dark to mid-green, opposite to alternate, basally more widely spaced than apically, often slightly overlapping, basal 2-4 pairs often gradually reduced, inaequilaterally lanceolate to trapezoid, acroscopically truncate, basiscopically excavate, usually alternately shallowly and deeply dentate, to 38 mm long, to 10 mm wide, glabrous adaxially and abaxially; costa adaxially raised, convex. Venation anadromous, evident, pinnately branched, branches forked, ending in the teeth near the margin. Sori linear, usually at or above a vein fork, to 4 mm long; indusium chartaceus, stramineous, transversely narrowly elliptic, entire, to 4 mm long, to 1.2 mm wide; sporangium long-stalked, uniseriate, 3-seriate below the capsule, capsule globose in lateral view, annulus with (19-)19(-22) indurated annulus cells, epistomium (2-)2(-3)-celled, hypostomium (4-)4(-5)-celled. Spores 64 per sporangium, brown, elliptic, monolete, perispore forms broad reticulate wings, erose, the areas between the wings irregularly echinulate, exospore (30-)32.2(-36) x (20-)22.3(-24) (m. Figure 48C & D.
Habitat
Ecology: Epilithic and restricted to moist, deeply shaded forests where the plants generally grow on wet or seasonally wet moss-covered rocks in or near perennial steams. Not edaphically bound, but in Swaziland the species is only known from the greenstone belt. Nanophanerophyte, mesophyte; fronds mesomorphic. Vegetative reproduction by the closely branched rhizome and rootbud formation. Seasonal pattern apparently nonexistent, no dormant period.
Range
Distribution: Frequent in the north-western corner of Swaziland, occurring at altitudes ranging between 500 and 1 250 m. The species is widespread is sub-Saharan Africa and the western Indian Ocean region.
Date Updated: 19 August 2007
Herbarium
South African National Biodiversity Institute, Compton Herbarium, Cape Town (SAM)
Collection
Swaziland ferns and fern allies
Resource Type
Reference Sources
Entry From
Swaziland Ferns and Fern Allies, (2003) Author: J.P. Roux
Names
Asplenium inaequilaterale Willd. [family ASPLENIACEAE]
Common names
Asplenium brachyotus Kunze in Linnaea 10: 512 (1836). Asplenium erectum Bory ex Willd. var. brachyotus (Kunze) Sim, Ferns S. Afr.: 138, LXVI, t. 1 (1892). Asplenium laetum Sw. var. brachyotus (Kunze) Bonap., Notes ptérid.16: 60 (1925). Type: Ad catarractum magnam in praeruptis umbrosis inter Omsamcaba, 200 m, Drège s.n. [LZ†, B!, lecto., designated by Roux (1986); BM!, K!, isolecto.]. Asplenium laetum sensu Sim, Ferns S. Afr., 2nd edn: 150, pl. 50. t. 1 (1915), non Sw. (1806).
inaequilateralis = with unequal sides
Vernacular names: Lolly spleenwort; Wortelknop tralievaring (Afr.)
Information
Plants epilithic. Rhizome short, erect to suberect, to 20 mm long, to 4 mm in diameter, set with roots, crowded persistent stipe bases and scales, scales crustaceous, atrocastaneus to black, clathrate, sessile, subulate to lanceolate, cordate to cordate-imbricate, entire or with scattered, pluricellular recurved outgrowths along the margin, apex terminates in a thin-walled cell, to 5 mm long, to 0.7 mm wide. Fronds crowded, caespitose, erect, to 430 mm long; stipe firm, castaneus, matt, terete, to 210 mm long, to 1.6 mm in diameter, narrowly green-winged to the base, base sparsely set with scales similar to those on the rhizome, higher up with filiform clathrate scales terminating in an elliptic thin-walled cell, to 1 mm long, and 3-4-celled hairs terminating in an enlarged obovate thin-walled cell, to 0.3 mm long; lamina 1-pinnate, narrowly elliptic, to 230 mm long, to 68 mm wide, with up to 17 petiolated pinna pairs; rachis firm, castaneus to greenish higher up, shallowly sulcate adaxially, narrowly green-winged, set with filiform scales and hairs similar to those on the stipe, glabrous later; pinnae basal pinnae petiolate, petiole to 1.5 mm long, sessile towards the apex, herbaceous, dark to mid-green, opposite to alternate, basally more widely spaced than apically, often slightly overlapping, basal 2-4 pairs often gradually reduced, inaequilaterally lanceolate to trapezoid, acroscopically truncate, basiscopically excavate, usually alternately shallowly and deeply dentate, to 38 mm long, to 10 mm wide, glabrous adaxially and abaxially; costa adaxially raised, convex. Venation anadromous, evident, pinnately branched, branches forked, ending in the teeth near the margin. Sori linear, usually at or above a vein fork, to 4 mm long; indusium chartaceus, stramineous, transversely narrowly elliptic, entire, to 4 mm long, to 1.2 mm wide; sporangium long-stalked, uniseriate, 3-seriate below the capsule, capsule globose in lateral view, annulus with (19-)19(-22) indurated annulus cells, epistomium (2-)2(-3)-celled, hypostomium (4-)4(-5)-celled. Spores 64 per sporangium, brown, elliptic, monolete, perispore forms broad reticulate wings, erose, the areas between the wings irregularly echinulate, exospore (30-)32.2(-36) x (20-)22.3(-24) (m. Figure 48C & D.
Habitat
Ecology: Epilithic and restricted to moist, deeply shaded forests where the plants generally grow on wet or seasonally wet moss-covered rocks in or near perennial steams. Not edaphically bound, but in Swaziland the species is only known from the greenstone belt. Nanophanerophyte, mesophyte; fronds mesomorphic. Vegetative reproduction by the closely branched rhizome and rootbud formation. Seasonal pattern apparently nonexistent, no dormant period.
Range
Distribution: Frequent in the north-western corner of Swaziland, occurring at altitudes ranging between 500 and 1 250 m. The species is widespread is sub-Saharan Africa and the western Indian Ocean region.
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