Information

Plants terrestrial or epilithic. Rhizome erect to suberect, to 35 mm long, to 3 mm in diameter, set with roots, crowded persistent stipe bases and scales, scales chartaceus to crustaceous, castaneus, clathrate, sessile, lanceolate, cordate to cordate-imbricate, with scattered filiform outgrowths along the margin of which the apex terminates in an elliptic thin-walled cell, apex terminates in an oblong thin-walled cell, to 8 mm long, to 1 mm wide. Fronds crowded, caespitose, erect to arching, to 510 mm long; stipe firm, castaneus, matt, terete, to 120 mm long, to 1.8 mm in diameter, narrowly green-winged in the upper half, initially moderately set with ferrugineous glandular cells and filiform clathrate scales of which the apex terminates in an enlarged gland-like cell, to 3 mm long, glabrous later; lamina 1-pinnate, narrowly elliptic to oblong-cuneate, to 390 mm long, to 60 mm wide, with up to 36 stalked pinna pairs; rachis firm, castaneus, terete, narrowly green-winged, initially moderately set with scales similar to those on the stipe, glabrous later; pinnae sessile, herbaceous, opposite to alternate, basally more widely spaced than apically, pinnae in the lower half to one third of the lamina gradually reduced, inaequilaterally lanceolate to ovate, often slightly auricled acroscopically, acroscopically truncate to broadly cuneate, basiscopically narrowly to broadly cuneate, dentate, to 30 mm long, to 14 mm wide, glabrous adaxially and abaxially; costa adaxially raised, convex. Venation anadromous, evident, pinnately branched, branches forked, ending in the teeth near the margin. Sori linear, inframedial, usually below or at a vein fork, to 3.5 mm long; indusium chartaceus, stramineous, linear, entire, to 3.5 mm long, to 0.3 mm wide; sporangium long-stalked, uniseriate, 3-seriate below the capsule, capsule globose in lateral view, annulus with (17-)19(-21) indurated annulus cells, epistomium 2-celled, hypostomium (3-)4(-4)-celled. Spores 64 per sporangium, brown, elliptic, monolete, perispore forms broad reticulate wings, exospore (26-)28.8(-38) x (16-)18.5(-24) (m. Figure 48A & B.