Entry From
Swaziland Ferns and Fern Allies, (2003) Author: J.P. Roux
Common names
Christella hilsenbergii (C.Presl) Holttum in J. S. African Bot. 40: 142 (1974); Jacobsen, Ferns Sthn. Afr.: 387, fig. 289 (1983). Nephrodium hilsenbergii C.Presl, Epimel. bot.: 47 (1851). Type: Mauritius, Sieber 49 (PRC, holo.; K, P, iso.). Nephrodium quadrangulare Fée, Mém. foug.: 308 (1852). Dryopteris quadrangularis (Fée) Alston in J. Bot. 75: 253 (1937). Cyclosorus quadrangularis (Fée) Tardieu in Notul. Syst. (Paris) 14: 345 (1953). Thelypteris quadrangularis (Fée) Schelpe in J. S. African Bot. 30: 196 (1964). Type: Habitat in Guyanâ gallicâ, in sylvis paludosis ad ripas amnis Mont-Tigre, Leprieur 182 (?P, holo.; NY iso.).
hispid = covered with rigid erect hairs
Vernacular names: Hispid marsh fern; Borselhaar moerasvaring (Afr.)
Information
Plants terrestrial. Rhizome suberect to erect, to 45 mm long, to 12 mm in diameter, densely set with roots, crowded persistent stipe bases and scales, scales chartaceus, castaneus, sessile, subulate, cordate, margins set with short filiform outgrowths, unicellular gland-like hairs and acicular hairs, apex terminates in a thin-walled cell, to 12 mm long, to 1.5 mm wide. Fronds crowded, erect to arching, to 970 mm long, to 13 per plant; stipe firm, stramineous, adaxially sulcate, to 425 mm long, to 3.5 mm in diameter, set with hairs and scales, hairs closely set, hyaline, acicular, to 0.3 mm long, scales usually restricted to the stipe base, similar to those on the rhizome; lamina 1-pinnate-pinnatifid, elliptic, to 540 mm long, to 250 mm wide, with up to 20 sessile pinna pairs; rachis stramineous to green, adaxially sulcate, sulcus not open to that of the costa, closely set with hyaline, unicellular acicular hairs, to 1.2 mm long; pinnae sessile, adnate towards the apex, herbaceous, oblong-cuneate, opposite to alternate, basally slightly more widely spaced, the 2 to 3 basal pinna pairs slightly and gradually reduced, to 140 mm long, to 17 mm wide, lobed, oblong-acute, falcate, crenulate, to 9 mm long, to 4 mm wide, adaxially with hyaline, unicellular acicular hairs along and between the veins, to 1.2 mm long, and with unicellular capitate glands on the lamina between the veins, to 80 (m long, abaxially set with hyaline, uni- and pluricellular acicular hairs along and between the veins, and with unicellular capitate glands on the lamina between the veins, to 80 (m long; costa adaxially sulcate, closely set with hyaline hairs similar to those on the rachis. Venation proximally anadromous, catadromous towards the apex, evident, pinnately branched in the segments, vein branches simple, with a single pair of veins anastomosing below the sinus, with the excurrent vein terminating in the sinus between the lobes. Sori circular, to 0.8 mm in diameter, medial on the veins, up to 8 pairs per segment; indusium firmly chartaceus, brown, persistent, reniform, to 0.5 mm in diameter, closely set with hyaline, unicellular acicular hairs, to 0.6 mm long; receptacle nude; sporangium long-stalked, 3-seriate below the capsule, usually with a single yellow-capped glandular cells, capsule broadly obovate in lateral view, with (13-)13(-15) indurated annulus cells, epistomium 3-celled, hypostomium (4-)4(-5)-celled. Spores 64 per sporangium, brown, elliptic to spheroidal, monolete, (28-)28.66(-34) x (18-)19.73(-22) (m. Figure 34C-E.