Information

Plants terrestrial. Rhizome wide-creeping, to 8 mm in diameter, irregularly branched, set with roots, spaced stipe bases, and sparsely set with scales, scales chartaceus, castaneus, sessile, ovate, cordate to cordate-imbricate, repand, often irregularly set with oblong thin-walled cells, apex terminates in an oblong thin-walled cell, to 2.5 mm long, to 1.5 mm wide. Fronds to 15 mm apart, straggling, to 1.3 mm long; stipe proximally black, brown to stramineous distally, adaxially shallowly sulcate, to 325 mm long, to 4 mm in diameter, set with hairs and scales, hairs are unicellular and acicular, to 0.3 mm long, scales sparsely set, chartaceus, ferrugineus, sessile, lanceolate to ovate, cordate-imbricate, margins variously set with oblong thin-walled cells, unicellular acicular cells and uniseriate hairs several cells long, of which the apical cell is enlarged, apex terminates in an oblong thin-walled cell, to 4.5 mm long, to 0.6 mm wide; lamina 1-pinnate, oblong-acute, to 1 m long, to 165 mm wide, with up to 22 free pinna pairs, the basal pinnae often significantly reduced, irregularly proliferous along the rachis; rachis stramineous, adaxially sulcate, sulcus not open to that of the costa, closely set with hairs similar to that on the stipe, also with 2-celled glandular hairs, sparsely scaled, scales chartaceus, ferrugineus to stramineous, sessile, ovate, cordate to cordate-imbricate, margins closely set with unicellular acicular hairs, apex terminates in an oblong thin-walled cell, to 1.5 mm long, to 0.4 mm wide, proliferating buds confined to the pinna axils, scaled, scales chartaceus, ferrugineus, similar to, but larger than, those on the rachis, to 1.5 mm long, to 0.5 mm wide; pinnae herbaceous, alternate, short-stalked, stalk to 1 mm long, oblong-acute, obtuse to truncate, often slightly inaequilateral, crenate, to 103 mm long, to 17 mm wide, adaxially glabrous, abaxially with hyaline, unicellular, acicular hairs along the costa and veins, sparsely scaled, scales chartaceus, ferrugineus, sessile, cordate, margins set with acicular hairs, apex terminates in an oblong cell, to 0.8 mm long, to 0.4 mm wide; costa adaxially shallowly sulcate, variously set with unicellular acicular hairs, abaxially pronounced, variously set with acicular hairs and scales similar to those on the rachis. Venation anadromous to catadromous, evident, pinnately branched in the lobes, veins simple, with up to 7 vein pairs of adjacent lobes anastomosing, the flexuose excurrent vein running to the sinus between two adjacent pinna lobes, the 2 to 5 free vein branch pairs end in the margin. Sori elongate, medial or inframedial on the anastomosing and free vein branches, sori often confluent and then without an excurrent vein, exindusiate; receptacle with simple, pluricellular hairs of which the apical cell is yellow and significantly enlarged; sporangium short-stalked, simple or occasionally with a hair similar to that on the receptacle, 3-seriate below the capsule, capsule globose to obovate, with (16-)18(-19) indurated annulus cells, epistomium (3-)4(-5)-celled, hypostomium (4-)4(-6)-celled. Spores 64 per sporangium, brown, ellipsoidal, monolete, echinulate. Figure 30C-E.

Information

Plants terrestrial. Rhizome wide-creeping, to 8 mm in diameter, irregularly branched, set with roots, spaced stipe bases, and sparsely set with scales, scales chartaceus, castaneus, sessile, ovate, cordate to cordate-imbricate, repand, often irregularly set with oblong thin-walled cells, apex terminates in an oblong thin-walled cell, to 2.5 mm long, to 1.5 mm wide. Fronds to 15 mm apart, straggling, to 1.3 mm long; stipe proximally black, brown to stramineous distally, adaxially shallowly sulcate, to 325 mm long, to 4 mm in diameter, set with hairs and scales, hairs are unicellular and acicular, to 0.3 mm long, scales sparsely set, chartaceus, ferrugineus, sessile, lanceolate to ovate, cordate-imbricate, margins variously set with oblong thin-walled cells, unicellular acicular cells and uniseriate hairs several cells long, of which the apical cell is enlarged, apex terminates in an oblong thin-walled cell, to 4.5 mm long, to 0.6 mm wide; lamina 1-pinnate, oblong-acute, to 1 m long, to 165 mm wide, with up to 22 free pinna pairs, the basal pinnae often significantly reduced, irregularly proliferous along the rachis; rachis stramineous, adaxially sulcate, sulcus not open to that of the costa, closely set with hairs similar to that on the stipe, also with 2-celled glandular hairs, sparsely scaled, scales chartaceus, ferrugineus to stramineous, sessile, ovate, cordate to cordate-imbricate, margins closely set with unicellular acicular hairs, apex terminates in an oblong thin-walled cell, to 1.5 mm long, to 0.4 mm wide, proliferating buds confined to the pinna axils, scaled, scales chartaceus, ferrugineus, similar to, but larger than, those on the rachis, to 1.5 mm long, to 0.5 mm wide; pinnae herbaceous, alternate, short-stalked, stalk to 1 mm long, oblong-acute, obtuse to truncate, often slightly inaequilateral, crenate, to 103 mm long, to 17 mm wide, adaxially glabrous, abaxially with hyaline, unicellular, acicular hairs along the costa and veins, sparsely scaled, scales chartaceus, ferrugineus, sessile, cordate, margins set with acicular hairs, apex terminates in an oblong cell, to 0.8 mm long, to 0.4 mm wide; costa adaxially shallowly sulcate, variously set with unicellular acicular hairs, abaxially pronounced, variously set with acicular hairs and scales similar to those on the rachis. Venation anadromous to catadromous, evident, pinnately branched in the lobes, veins simple, with up to 7 vein pairs of adjacent lobes anastomosing, the flexuose excurrent vein running to the sinus between two adjacent pinna lobes, the 2 to 5 free vein branch pairs end in the margin. Sori elongate, medial or inframedial on the anastomosing and free vein branches, sori often confluent and then without an excurrent vein, exindusiate; receptacle with simple, pluricellular hairs of which the apical cell is yellow and significantly enlarged; sporangium short-stalked, simple or occasionally with a hair similar to that on the receptacle, 3-seriate below the capsule, capsule globose to obovate, with (16-)18(-19) indurated annulus cells, epistomium (3-)4(-5)-celled, hypostomium (4-)4(-6)-celled. Spores 64 per sporangium, brown, ellipsoidal, monolete, echinulate. Figure 30C-E.