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Crassula nodulosa forma nodulosa [family CRASSULACEAE]
Date Updated: 26 July 2007
Herbarium
Royal Botanic Gardens, Kew (K)
Collection
Flora Zambesiaca
Resource Type
Reference Sources
Entry From
FZ, Vol 7, Part 1, page 3, (1983) Author: R. Fernandes
Names
Crassula nodulosa forma nodulosa [family CRASSULACEAE]
Habitat
In grassland, bushveld in rocky situations, in mountains
Range
perhaps also in Uganda, Kenya and Tanzania.
Altitude range
up to 2200 m.
2200
0
inferred only top
Distribution
Mozambique MS Manica e Sofala, c. 1440 m., 8.x.1969, Leach & Cannell 14397 (SRGH).Zimbabwe E Inyangani Mt., 2000-2200 m., 21-22.iv.1934, Humbert 15788 (P).Zimbabwe C Salisbury, Umvukwe Mt., 1680 m., 5.iii.1961, Richards 14576 (K).
Distribution (external)
Angola
Namibia
S. Africa (Transvaal, Orange Free State and the Cape)
Notes
Most of the synonyms given above under C. nodulosa apply to this taxon. The type of C. data, however, consists only of the upper part of the stem and thus it is impossible to tell the shape of the median leaves. C. nodulosa was treated by Tölken (loc. cit., 1975 & 1977) as a subspecies of C. capitella. We interpret it as an independent species differing from C. capitella as follows:Perennial rarely biennial, with a tuberous base and usually ± swollen root; stems hairy or papillose; mucro of petals solid, conical, usually acute, tuberculate-scabrid, inserted near the apex of petals, the latter inflexed and obtuse; young follicles at first shortly attenuate with stigmas on very short styles, the stigmas becoming lateral at maturity; nectary-scales truncate or slightly emarginate, with thickened upper margin; seeds minutely tuberculateC. nodulosaBiennial rarely perennial without tuberous base and with several main roots; stems glabrous; appendage of petals hollow, broader than above, subcylindrical, obtuse, smooth, inserted further from the apex of the petals, the latter more inflexed than above and acute; young follicles having sessile lateral stigmas; nectary scales distinctly and more deeply emarginate than above, not thickened; seeds smoothC. capitellaTölken placed plants with a tuberculate-scabrid petal-appendage in C. capitella subsp. enantiophylla, this includes all specimens from the F.Z. area. According to his keys, subsp. nodulosa has a smooth petal-mucro. However, not only the type of C. nodulosa (Adams 28), but also many plants from the Transvaal and the Cape Prov. determined by Tölken as subsp. nodulosa have distinctly tuberculate-scabrid mucros. He refers to the appendage as “terminal”, but this is inexact as the position is actually subterminal and only appears terminal by the inflexion of the petal apex. The type of C. enanthiophylla is a plant with laxer cymes than typical C. nodulosa, a condition which we think was probably related to its shady habitat. Many plants from the F.Z. area of var. nodulosa forma nodulosa have the calyx somewhat longer than is usual in material from other regions (Transvaal, Angola, etc.) thus being intermediate in this respect between var. nodulosa and var. longisepala.
Date Updated: 26 July 2007
Herbarium
Royal Botanic Gardens, Kew (K)
Collection
Flora Zambesiaca
Resource Type
Reference Sources
Entry From
FZ, Vol 7, Part 1, page 3, (1983) Author: R. Fernandes
Names
Crassula nodulosa forma nodulosa [family CRASSULACEAE]
Habitat
In grassland, bushveld in rocky situations, in mountains
Range
perhaps also in Uganda, Kenya and Tanzania.
Altitude range
up to 2200 m.
2200
0
inferred only top
Distribution
Mozambique MS Manica e Sofala, c. 1440 m., 8.x.1969, Leach & Cannell 14397 (SRGH).Zimbabwe E Inyangani Mt., 2000-2200 m., 21-22.iv.1934, Humbert 15788 (P).Zimbabwe C Salisbury, Umvukwe Mt., 1680 m., 5.iii.1961, Richards 14576 (K).
Distribution (external)
Angola
Namibia
S. Africa (Transvaal, Orange Free State and the Cape)
Notes
Most of the synonyms given above under C. nodulosa apply to this taxon. The type of C. data, however, consists only of the upper part of the stem and thus it is impossible to tell the shape of the median leaves. C. nodulosa was treated by Tölken (loc. cit., 1975 & 1977) as a subspecies of C. capitella. We interpret it as an independent species differing from C. capitella as follows:Perennial rarely biennial, with a tuberous base and usually ± swollen root; stems hairy or papillose; mucro of petals solid, conical, usually acute, tuberculate-scabrid, inserted near the apex of petals, the latter inflexed and obtuse; young follicles at first shortly attenuate with stigmas on very short styles, the stigmas becoming lateral at maturity; nectary-scales truncate or slightly emarginate, with thickened upper margin; seeds minutely tuberculateC. nodulosaBiennial rarely perennial without tuberous base and with several main roots; stems glabrous; appendage of petals hollow, broader than above, subcylindrical, obtuse, smooth, inserted further from the apex of the petals, the latter more inflexed than above and acute; young follicles having sessile lateral stigmas; nectary scales distinctly and more deeply emarginate than above, not thickened; seeds smoothC. capitellaTölken placed plants with a tuberculate-scabrid petal-appendage in C. capitella subsp. enantiophylla, this includes all specimens from the F.Z. area. According to his keys, subsp. nodulosa has a smooth petal-mucro. However, not only the type of C. nodulosa (Adams 28), but also many plants from the Transvaal and the Cape Prov. determined by Tölken as subsp. nodulosa have distinctly tuberculate-scabrid mucros. He refers to the appendage as “terminal”, but this is inexact as the position is actually subterminal and only appears terminal by the inflexion of the petal apex. The type of C. enanthiophylla is a plant with laxer cymes than typical C. nodulosa, a condition which we think was probably related to its shady habitat. Many plants from the F.Z. area of var. nodulosa forma nodulosa have the calyx somewhat longer than is usual in material from other regions (Transvaal, Angola, etc.) thus being intermediate in this respect between var. nodulosa and var. longisepala.
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