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ALCHEMILLA kiwuensis Engl. [family ROSACEAE]

Date Updated: 19 August 2007

Herbarium
Royal Botanic Gardens, Kew (K)
Collection
Flora of Tropical East Africa
Resource Type
Reference Sources
Entry From
Flora of Tropical East Africa, page 1, (1960) Author: R. A. GRAHAM
Names
ALCHEMILLA kiwuensis Engl. [family ROSACEAE], in Z.A.E.: 225, t. 21 F–G (1911); Hauman & Balle in Rev. Zool. Bot. Afr. 24: 308 (1934); F.P.N.A. 1: 244, t. 23 (1948); F.C.B. 3: 6, t. 1 (1952); Hedb. in Symb. Bot. Ups. 15 (1): 102 (1957); F.W.T.A., ed. 2, 1: 424 (1958). Type: Ruanda-Urundi, Lake Kalago, Mildbraed 1538 (B, holo.†, BR, fragment, ? holo.!) ALCHEMILLA adolphi-friedericii Engl. [family ROSACEAE], in Z.A.E.: 225, t. 21 A–E (1911). Type: Ruanda-Urundi, Rukarara, Mildbraed 889 (B, holo. †) ALCHEMILLA kiwuensis Engl. var. kandtiana [family ROSACEAE], in Z.A.E.: 225 (1911). Type: Ruanda-Urundi, Mt. Niansa, Kandt 123 (B, holo. †)
Information
A low herb usually with basal rosettes with or without creeping, ± slender runners covered with long spreading white hairs, often rooting and developing secondary rosettes at intervals. Stipules commonly 1–2.5 cm. long, united below, free and foliaceous above, the dorsal and ventral sutures being nearly of equal length, thus the aperture scarcely oblique. Rosette-leaves petiolate, rounded to reniform in outline, 5–7(–9)-lobed, 1.8–3(–4.5) cm. long, 2.5–4.5(–7) cm. wide, variably hairy on both surfaces with whitish hairs; lobes usually reaching beyond the centre of the lamina, oblong-elliptic to obovate, terminally rounded, the teeth up to 2 mm. deep and continuing down the sides of the lobes which are entire only towards the base; petiole 3–12 cm. long. Stem-leaves smaller, usually 5-lobed, 1–1.5 cm. long, 1.5–2.5 cm. wide; petiole 1.5–2.5 cm. long. Flowers in 1–4-flowered cymose clusters or in shortly branched panicles; the inflorescence rarely of only 1–2 flowers hidden by the stipules as in A. cryptantha; pedicels short, up to 1 mm. long, glabrous or hairy. Calyx-tube ± 2 mm. long, covered with long, spreading white hairs; calyx-lobes 0.75 mm. long, ovate-triangular; epicalyx-lobes exceeding the calyx-lobes, 1–1.25 mm. long, broadly ovate. Achenes 5–8.
Range
DISTR. U1–3; K3, 4; T2–4, 6–8
Altitude range
1250–3000 m.
Distribution
KENYA Ravine District Eldama Ravine, Whyte!KENYA Aberdare Mts., Kinangop, Sept. 1951, Verdcourt 601!KENYA Mt. Kenya, western side, Fries 613A!, 613B!TANGANYIKA Moshi District Lyamungu, Oct. 1943, Wallace 1099B!;TANGANYIKA Ufipa District Malonje, Apr. 1950, Buttock 2780!;TANGANYIKA Rungwe District Kiwira R., Lower Fishing Camp, Oct. 1947, Greenway & Brenan 8264!UGANDA Karamoja District Mt. Debasien, Jan. 1936, Eggeling 2698!;UGANDA Kigezi District Behungi, Dec. 1933, A. S. Thomas 1159!UGANDA Ruwenzori, Aug. 1938, Purseglove 298!
Distribution (external)
; Fernando Po British Cameroons the Belgian Congo Southern Rhodesia
Notes
This species is distinguished from A. cryptantha by the longer, laterally more dentate leaf-lobes, by the tendency to produce persistent primary and secondary rosettes, by the stipules being subequally sutured, and by the stronger, cymose or paniculate inflorescences. There is also an apparent tendency for the urceoles of A. cryptantha to be glabrous, whereas in A. kiwuensis they are covered with long hairs. Conversely the pedicels of the latter are sometimes glabrous, a feature not noticed in A. cryptantha. But none of these characters provides a clear distinction, there being many intermediates, due perhaps either to overlapping of the limits of variation of each or to interspecific hybridity. Bullock 2780 approaches subsp. rhodesica Hauman & Balle (Rev. Zool. Bot. Afr. 24: 342 (1934)) in having some densely hairy petioles and subglabrous flowers with calyx-lobes nearly equalling the epicalyx, but examples corresponding exactly to the description of this subspecies, which does not seem to be very clearly distinguishable, have not yet been found in our area.

Date Updated: 19 August 2007

Herbarium
Royal Botanic Gardens, Kew (K)
Collection
Flora of Tropical East Africa
Resource Type
Reference Sources
Entry From
Flora of Tropical East Africa, page 1, (1960) Author: R. A. GRAHAM
Names
ALCHEMILLA kiwuensis Engl. [family ROSACEAE], in Z.A.E.: 225, t. 21 F–G (1911); Hauman & Balle in Rev. Zool. Bot. Afr. 24: 308 (1934); F.P.N.A. 1: 244, t. 23 (1948); F.C.B. 3: 6, t. 1 (1952); Hedb. in Symb. Bot. Ups. 15 (1): 102 (1957); F.W.T.A., ed. 2, 1: 424 (1958). Type: Ruanda-Urundi, Lake Kalago, Mildbraed 1538 (B, holo.†, BR, fragment, ? holo.!) ALCHEMILLA adolphi-friedericii Engl. [family ROSACEAE], in Z.A.E.: 225, t. 21 A–E (1911). Type: Ruanda-Urundi, Rukarara, Mildbraed 889 (B, holo. †) ALCHEMILLA kiwuensis Engl. var. kandtiana [family ROSACEAE], in Z.A.E.: 225 (1911). Type: Ruanda-Urundi, Mt. Niansa, Kandt 123 (B, holo. †)
Information
A low herb usually with basal rosettes with or without creeping, ± slender runners covered with long spreading white hairs, often rooting and developing secondary rosettes at intervals. Stipules commonly 1–2.5 cm. long, united below, free and foliaceous above, the dorsal and ventral sutures being nearly of equal length, thus the aperture scarcely oblique. Rosette-leaves petiolate, rounded to reniform in outline, 5–7(–9)-lobed, 1.8–3(–4.5) cm. long, 2.5–4.5(–7) cm. wide, variably hairy on both surfaces with whitish hairs; lobes usually reaching beyond the centre of the lamina, oblong-elliptic to obovate, terminally rounded, the teeth up to 2 mm. deep and continuing down the sides of the lobes which are entire only towards the base; petiole 3–12 cm. long. Stem-leaves smaller, usually 5-lobed, 1–1.5 cm. long, 1.5–2.5 cm. wide; petiole 1.5–2.5 cm. long. Flowers in 1–4-flowered cymose clusters or in shortly branched panicles; the inflorescence rarely of only 1–2 flowers hidden by the stipules as in A. cryptantha; pedicels short, up to 1 mm. long, glabrous or hairy. Calyx-tube ± 2 mm. long, covered with long, spreading white hairs; calyx-lobes 0.75 mm. long, ovate-triangular; epicalyx-lobes exceeding the calyx-lobes, 1–1.25 mm. long, broadly ovate. Achenes 5–8.
Range
DISTR. U1–3; K3, 4; T2–4, 6–8
Altitude range
1250–3000 m.
Distribution
KENYA Ravine District Eldama Ravine, Whyte!KENYA Aberdare Mts., Kinangop, Sept. 1951, Verdcourt 601!KENYA Mt. Kenya, western side, Fries 613A!, 613B!TANGANYIKA Moshi District Lyamungu, Oct. 1943, Wallace 1099B!;TANGANYIKA Ufipa District Malonje, Apr. 1950, Buttock 2780!;TANGANYIKA Rungwe District Kiwira R., Lower Fishing Camp, Oct. 1947, Greenway & Brenan 8264!UGANDA Karamoja District Mt. Debasien, Jan. 1936, Eggeling 2698!;UGANDA Kigezi District Behungi, Dec. 1933, A. S. Thomas 1159!UGANDA Ruwenzori, Aug. 1938, Purseglove 298!
Distribution (external)
; Fernando Po British Cameroons the Belgian Congo Southern Rhodesia
Notes
This species is distinguished from A. cryptantha by the longer, laterally more dentate leaf-lobes, by the tendency to produce persistent primary and secondary rosettes, by the stipules being subequally sutured, and by the stronger, cymose or paniculate inflorescences. There is also an apparent tendency for the urceoles of A. cryptantha to be glabrous, whereas in A. kiwuensis they are covered with long hairs. Conversely the pedicels of the latter are sometimes glabrous, a feature not noticed in A. cryptantha. But none of these characters provides a clear distinction, there being many intermediates, due perhaps either to overlapping of the limits of variation of each or to interspecific hybridity. Bullock 2780 approaches subsp. rhodesica Hauman & Balle (Rev. Zool. Bot. Afr. 24: 342 (1934)) in having some densely hairy petioles and subglabrous flowers with calyx-lobes nearly equalling the epicalyx, but examples corresponding exactly to the description of this subspecies, which does not seem to be very clearly distinguishable, have not yet been found in our area.

Date Updated: 19 August 2007

Herbarium
Royal Botanic Gardens, Kew (K)
Collection
Flora of Tropical East Africa
Resource Type
Reference Sources
Entry From
Flora of Tropical East Africa, page 1, (1960) Author: R. A. GRAHAM
Names
ALCHEMILLA kiwuensis Engl. [family ROSACEAE], in Z.A.E.: 225, t. 21 F–G (1911); Hauman & Balle in Rev. Zool. Bot. Afr. 24: 308 (1934); F.P.N.A. 1: 244, t. 23 (1948); F.C.B. 3: 6, t. 1 (1952); Hedb. in Symb. Bot. Ups. 15 (1): 102 (1957); F.W.T.A., ed. 2, 1: 424 (1958). Type: Ruanda-Urundi, Lake Kalago, Mildbraed 1538 (B, holo.†, BR, fragment, ? holo.!) ALCHEMILLA adolphi-friedericii Engl. [family ROSACEAE], in Z.A.E.: 225, t. 21 A–E (1911). Type: Ruanda-Urundi, Rukarara, Mildbraed 889 (B, holo. †) ALCHEMILLA kiwuensis Engl. var. kandtiana [family ROSACEAE], in Z.A.E.: 225 (1911). Type: Ruanda-Urundi, Mt. Niansa, Kandt 123 (B, holo. †)
Information
A low herb usually with basal rosettes with or without creeping, ± slender runners covered with long spreading white hairs, often rooting and developing secondary rosettes at intervals. Stipules commonly 1–2.5 cm. long, united below, free and foliaceous above, the dorsal and ventral sutures being nearly of equal length, thus the aperture scarcely oblique. Rosette-leaves petiolate, rounded to reniform in outline, 5–7(–9)-lobed, 1.8–3(–4.5) cm. long, 2.5–4.5(–7) cm. wide, variably hairy on both surfaces with whitish hairs; lobes usually reaching beyond the centre of the lamina, oblong-elliptic to obovate, terminally rounded, the teeth up to 2 mm. deep and continuing down the sides of the lobes which are entire only towards the base; petiole 3–12 cm. long. Stem-leaves smaller, usually 5-lobed, 1–1.5 cm. long, 1.5–2.5 cm. wide; petiole 1.5–2.5 cm. long. Flowers in 1–4-flowered cymose clusters or in shortly branched panicles; the inflorescence rarely of only 1–2 flowers hidden by the stipules as in A. cryptantha; pedicels short, up to 1 mm. long, glabrous or hairy. Calyx-tube ± 2 mm. long, covered with long, spreading white hairs; calyx-lobes 0.75 mm. long, ovate-triangular; epicalyx-lobes exceeding the calyx-lobes, 1–1.25 mm. long, broadly ovate. Achenes 5–8.
Range
DISTR. U1–3; K3, 4; T2–4, 6–8
Altitude range
1250–3000 m.
Distribution
KENYA Ravine District Eldama Ravine, Whyte!KENYA Aberdare Mts., Kinangop, Sept. 1951, Verdcourt 601!KENYA Mt. Kenya, western side, Fries 613A!, 613B!TANGANYIKA Moshi District Lyamungu, Oct. 1943, Wallace 1099B!;TANGANYIKA Ufipa District Malonje, Apr. 1950, Buttock 2780!;TANGANYIKA Rungwe District Kiwira R., Lower Fishing Camp, Oct. 1947, Greenway & Brenan 8264!UGANDA Karamoja District Mt. Debasien, Jan. 1936, Eggeling 2698!;UGANDA Kigezi District Behungi, Dec. 1933, A. S. Thomas 1159!UGANDA Ruwenzori, Aug. 1938, Purseglove 298!
Distribution (external)
; Fernando Po British Cameroons the Belgian Congo Southern Rhodesia
Notes
This species is distinguished from A. cryptantha by the longer, laterally more dentate leaf-lobes, by the tendency to produce persistent primary and secondary rosettes, by the stipules being subequally sutured, and by the stronger, cymose or paniculate inflorescences. There is also an apparent tendency for the urceoles of A. cryptantha to be glabrous, whereas in A. kiwuensis they are covered with long hairs. Conversely the pedicels of the latter are sometimes glabrous, a feature not noticed in A. cryptantha. But none of these characters provides a clear distinction, there being many intermediates, due perhaps either to overlapping of the limits of variation of each or to interspecific hybridity. Bullock 2780 approaches subsp. rhodesica Hauman & Balle (Rev. Zool. Bot. Afr. 24: 342 (1934)) in having some densely hairy petioles and subglabrous flowers with calyx-lobes nearly equalling the epicalyx, but examples corresponding exactly to the description of this subspecies, which does not seem to be very clearly distinguishable, have not yet been found in our area.
ALCHEMILLA kiwuensis Engl. [family ROSACEAE]
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