Entry From
Flora of South Africa, (2003) Author: Dr J.P. Roux
Information
Erect dense twiggy evergreen shrubs, 0-75-5m high or trees up to 7 m high with a densely branched canopy, stems or trunks 2-15 cm in diam., bark grey, scaly; branchlets much divided, densely covered with leaves, glabrous except for rust-coloured glands which often cover the young parts giving the surface a rust-brown granular appearance. Leaves subopposite, alternate or in pseudo-whorls at ends of the branches, coriaceous (in texture often resembling old leather), shortly petioled, pale to dark green, paler and often rust-brown below when young, quite glabrous but sparsely to densely covered with rust-coloured glands which give the surface a granular appearance, obovate, oblanceolate, narrowly-elliptic or narrowly oblanceolate; base cuneate; apex obtuse, subobtuse or abruptly narrowed to a rounded tip; nervation indistinct on both surfaces or raised on upper surface, usually not raised on lower surface; margin entire, not revolute, very strongly undulate to practically flat; petioles 1-3 mm long. Inflorescence 5-20 mm long, axillary, glabrous except for rust-coloured, often stalked glands, 5-7-flowered; bracts lanceolate, boat-shaped, glandular. Flowers dioecious, white, greenish-white or cream, fragrant, mostly tetramerous. Male flowers somewhat larger than the females, 3-3 • 5 mm long. Calyx shallowly saucer-shaped, shallowly 4-(5-6-)lobed, glandular on the outside, \ or less of the length of the corolla. Corolla cup-shaped to widely bell-shaped, glabrous or with a few hairs on the lobes; tube cleft halfway or more; lobes 4(5-6), ovate. Stamens 12-20, in 2 rows, often in pairs together; filaments glabrous; anthers lanceolate to oblong-lanceolate, hairy at the apex. Ovary rudimentary, situated on a fringed disc, occasionally with 2 glabrous styles and then densely covered with whitish glands. Female flowers smaller than the males, 2-2 • 5 mm long. Calyx saucer-shaped 4-(5-6-)lobed on the rim, glandular outside. Corolla usually campanulate, tube cleft halfway down or more, lobes 4(5-6) usually contorted, as long as or longer than the tube, usually glabrous, occasionally with a few appressed bristles. Staminodes present or absent, star-shaped, hairy upwards. Ovary depressed ovoid, situated on a fringed disc, usually densely covered with white or grey scale-like glands, usually 2-celled due to incomplete septation of two cells with septa present only in the form of a ridge (4-6-celled ovaries also found with the cells complete and a single pendulous ovule in each); styles usually 2, stout, joined at the base, glabrous; stigma obliquely crescent-shaped. Fruit a globose, fleshy berry, 4-6 mm in diam., one-seeded, red, turning purple or black when ripe. Seeds globose, 3-4 mm in diam., divided into three parts by two thin curved lines and a groove; endosperm flinty, pale grey.
Use
12. Euclea undulata Thunb., Prodr. 85 (1796). Type: Cape of Good Hope, Thunberg s.n. (UPS, holo.!).
Range
A very widespread species occurring in the Cape, South West Africa, Bechuanaland, western and northern Transvaal, and the lowveld areas of northern Natal and Swaziland. It occurs sporadically in Southern Rhodesia but this seems to be the northernmost limit of its distribution and except for a number of specimens from Bechuanaland it has not been recorded from other parts of tropical Africa. This species often closely resembles E. divinorum but differs from it in the glandular not hairy ovary, the smaller leaves and fruits, the usually glabrous corolla and the less dense not compound inflorescence. The vernacular name used for both varieties is Guarrie or Gwarrie. The fruits have been reported as edible in both varieties but are not very palatable. The leaves of var. undulata as well as var. myrtina are reported to be browsed by cattle and the var. myrtina by kudu in South West Africa.