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COMMIPHORA Jacq. [family BURSERACEAE]
Date Updated: 19 August 2007
Herbarium
Royal Botanic Gardens, Kew (K)
Collection
Flora Somalia
Resource Type
Reference Sources
Entry From
Flora Somalia, Vol 2, (1999) Author: by M. Thulin [updated by M. Thulin 2008]
Names
COMMIPHORA Jacq. [family BURSERACEAE], (1797)
Information
Trees or shrubs, dioecious or rarely monoecious; outer bark often peeling away from the green underbark; exudate usually aromatic, often drying to a gum-resin; wood soft, light; branches often spine-tipped. Leaves imparipinnate, 1–3-foliolate, or simple. Flowers usually produced before the leaves, in panicles, cymes, clusters or solitary, the male inflorescences often larger and looser than the female. Calyx 4-lobed. Petals 4, free. Stamens 8 in two whorls of 4, or sometimes 4; staminodes present in female flowers. Ovary 2(–3)-celled, with 2 pendulous ovules in each cell; style short; stigma capitate; rudimentary ovary sometimes present in male flowers. Fruit a drupe with a single 1(–2)-seeded stone; outer part of pericarp (fruit wall) splitting into 2 or 4 valves; stone usually covered, at least at the base, by a ± fleshy, variously lobed, and often red, orange or yellow pseudaril (an outgrowth of the endocarp), usually with a fertile 1-seeded cell and a sterile cell, but sometimes both cells fertile.
Range
Genus of some 150 species, widespread in the drier parts of tropical Africa and Madagascar and extending to India via the Arabian peninsula and Iran, and in Mexico and Brazil.
Notes
Apart from being dioecious, many species of Commiphora are leafless for most of the year and usually set flower and fruit when leafless or with young leaves. Therefore, the collecting of complete material is difficult, and many species have been described from incomplete or mixed collections. This account owes much to the work done by the late Jan B. Gillett at Kew, although many of the taxa preliminary proposed by him have not been accepted. Kaj Vollesen’s account of Commiphora in Fl. Eth. 3 (1990) has also been very useful. Many of the species are still incompletely known and further good material is often needed to improve the descriptions. Even in cases when a flowering or fruiting tree looks leafless, dried up leaves are often present and should be collected. Information on bark characteristics and presence/absence of spines is essential. For identification by the key below branches with leaves and ripe fruits are usually needed. To count the number of valves of the fruits and to see the pseudaril in herbarium material it is often necessary to soften the fruit by boiling it for some time. In most cases, such as in the species of sect. Opobalsamae (see below), it is usually possible to see that the fruits are 4-valved already at an early stage, as they are marked with four longitudinal pale lines. In other species, such as C. playfairii, young fruits appear 2-valved and it is necessary to have ripe fruits to see that the valves actually partially split again. The species of Commiphora are an extremely important constituent of the vegetation in Somalia and dominate over large areas, usually along with species of Acacia. They are important as browse for the livestock, and several species also produce valuable gum-resins. The most important of these is myrrh, produced by C. myrrha. The genus Commiphora has been divided into a number of more or less well marked sections. Following the system employed by Gillett in Fl. Trop. E. Afr. (1991) the Somali species belong to the following sections: sect. Arillopsidium (species 1–8), sect. Hemprichia (species 9–12), sect. Coriacea (species 13), sect. Africanae (species 14–17), sect. Hildebrandtianae (species 18–25), sect. Rostratae (species 26), sect. Ciliatae (species 27), sect. Latifoliolatae (species 28–30), sect. Campestres (species 31–34), sect. Commiphora (species 35–40), sect. Opobalsamae (species 41–50). Species 51–53 cannot be placed in a section with certainty at present.
Date Updated: 19 August 2007
Herbarium
Royal Botanic Gardens, Kew (K)
Collection
Flora Somalia
Resource Type
Reference Sources
Entry From
Flora Somalia, Vol 2, (1999) Author: by M. Thulin [updated by M. Thulin 2008]
Names
COMMIPHORA Jacq. [family BURSERACEAE], (1797)
Information
Trees or shrubs, dioecious or rarely monoecious; outer bark often peeling away from the green underbark; exudate usually aromatic, often drying to a gum-resin; wood soft, light; branches often spine-tipped. Leaves imparipinnate, 1–3-foliolate, or simple. Flowers usually produced before the leaves, in panicles, cymes, clusters or solitary, the male inflorescences often larger and looser than the female. Calyx 4-lobed. Petals 4, free. Stamens 8 in two whorls of 4, or sometimes 4; staminodes present in female flowers. Ovary 2(–3)-celled, with 2 pendulous ovules in each cell; style short; stigma capitate; rudimentary ovary sometimes present in male flowers. Fruit a drupe with a single 1(–2)-seeded stone; outer part of pericarp (fruit wall) splitting into 2 or 4 valves; stone usually covered, at least at the base, by a ± fleshy, variously lobed, and often red, orange or yellow pseudaril (an outgrowth of the endocarp), usually with a fertile 1-seeded cell and a sterile cell, but sometimes both cells fertile.
Range
Genus of some 150 species, widespread in the drier parts of tropical Africa and Madagascar and extending to India via the Arabian peninsula and Iran, and in Mexico and Brazil.
Notes
Apart from being dioecious, many species of Commiphora are leafless for most of the year and usually set flower and fruit when leafless or with young leaves. Therefore, the collecting of complete material is difficult, and many species have been described from incomplete or mixed collections. This account owes much to the work done by the late Jan B. Gillett at Kew, although many of the taxa preliminary proposed by him have not been accepted. Kaj Vollesen’s account of Commiphora in Fl. Eth. 3 (1990) has also been very useful. Many of the species are still incompletely known and further good material is often needed to improve the descriptions. Even in cases when a flowering or fruiting tree looks leafless, dried up leaves are often present and should be collected. Information on bark characteristics and presence/absence of spines is essential. For identification by the key below branches with leaves and ripe fruits are usually needed. To count the number of valves of the fruits and to see the pseudaril in herbarium material it is often necessary to soften the fruit by boiling it for some time. In most cases, such as in the species of sect. Opobalsamae (see below), it is usually possible to see that the fruits are 4-valved already at an early stage, as they are marked with four longitudinal pale lines. In other species, such as C. playfairii, young fruits appear 2-valved and it is necessary to have ripe fruits to see that the valves actually partially split again. The species of Commiphora are an extremely important constituent of the vegetation in Somalia and dominate over large areas, usually along with species of Acacia. They are important as browse for the livestock, and several species also produce valuable gum-resins. The most important of these is myrrh, produced by C. myrrha. The genus Commiphora has been divided into a number of more or less well marked sections. Following the system employed by Gillett in Fl. Trop. E. Afr. (1991) the Somali species belong to the following sections: sect. Arillopsidium (species 1–8), sect. Hemprichia (species 9–12), sect. Coriacea (species 13), sect. Africanae (species 14–17), sect. Hildebrandtianae (species 18–25), sect. Rostratae (species 26), sect. Ciliatae (species 27), sect. Latifoliolatae (species 28–30), sect. Campestres (species 31–34), sect. Commiphora (species 35–40), sect. Opobalsamae (species 41–50). Species 51–53 cannot be placed in a section with certainty at present.
Date Updated: 19 August 2007
Herbarium
Royal Botanic Gardens, Kew (K)
Collection
Flora Somalia
Resource Type
Reference Sources
Entry From
Flora Somalia, Vol 2, (1999) Author: by M. Thulin [updated by M. Thulin 2008]
Names
COMMIPHORA Jacq. [family BURSERACEAE], (1797)
Information
Trees or shrubs, dioecious or rarely monoecious; outer bark often peeling away from the green underbark; exudate usually aromatic, often drying to a gum-resin; wood soft, light; branches often spine-tipped. Leaves imparipinnate, 1–3-foliolate, or simple. Flowers usually produced before the leaves, in panicles, cymes, clusters or solitary, the male inflorescences often larger and looser than the female. Calyx 4-lobed. Petals 4, free. Stamens 8 in two whorls of 4, or sometimes 4; staminodes present in female flowers. Ovary 2(–3)-celled, with 2 pendulous ovules in each cell; style short; stigma capitate; rudimentary ovary sometimes present in male flowers. Fruit a drupe with a single 1(–2)-seeded stone; outer part of pericarp (fruit wall) splitting into 2 or 4 valves; stone usually covered, at least at the base, by a ± fleshy, variously lobed, and often red, orange or yellow pseudaril (an outgrowth of the endocarp), usually with a fertile 1-seeded cell and a sterile cell, but sometimes both cells fertile.
Range
Genus of some 150 species, widespread in the drier parts of tropical Africa and Madagascar and extending to India via the Arabian peninsula and Iran, and in Mexico and Brazil.
Notes
Apart from being dioecious, many species of Commiphora are leafless for most of the year and usually set flower and fruit when leafless or with young leaves. Therefore, the collecting of complete material is difficult, and many species have been described from incomplete or mixed collections. This account owes much to the work done by the late Jan B. Gillett at Kew, although many of the taxa preliminary proposed by him have not been accepted. Kaj Vollesen’s account of Commiphora in Fl. Eth. 3 (1990) has also been very useful. Many of the species are still incompletely known and further good material is often needed to improve the descriptions. Even in cases when a flowering or fruiting tree looks leafless, dried up leaves are often present and should be collected. Information on bark characteristics and presence/absence of spines is essential. For identification by the key below branches with leaves and ripe fruits are usually needed. To count the number of valves of the fruits and to see the pseudaril in herbarium material it is often necessary to soften the fruit by boiling it for some time. In most cases, such as in the species of sect. Opobalsamae (see below), it is usually possible to see that the fruits are 4-valved already at an early stage, as they are marked with four longitudinal pale lines. In other species, such as C. playfairii, young fruits appear 2-valved and it is necessary to have ripe fruits to see that the valves actually partially split again. The species of Commiphora are an extremely important constituent of the vegetation in Somalia and dominate over large areas, usually along with species of Acacia. They are important as browse for the livestock, and several species also produce valuable gum-resins. The most important of these is myrrh, produced by C. myrrha. The genus Commiphora has been divided into a number of more or less well marked sections. Following the system employed by Gillett in Fl. Trop. E. Afr. (1991) the Somali species belong to the following sections: sect. Arillopsidium (species 1–8), sect. Hemprichia (species 9–12), sect. Coriacea (species 13), sect. Africanae (species 14–17), sect. Hildebrandtianae (species 18–25), sect. Rostratae (species 26), sect. Ciliatae (species 27), sect. Latifoliolatae (species 28–30), sect. Campestres (species 31–34), sect. Commiphora (species 35–40), sect. Opobalsamae (species 41–50). Species 51–53 cannot be placed in a section with certainty at present.
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