Plants terrestrial or epilithic. Rhizome short-decumbent, closely branched, to 2 mm in diameter, closely set with roots, persistent stipe bases and bicolorous scales, scales with a crustaceous, black to castaneus central stripe, nitid, and ferrugineus to stramineous, chartaceus to membranous margins, sessile, linear to narrowly lanceolate, base somewhat round-auricled, cordate, entire to repand, apex terminates in an oblong thin-walled cell, to 4 mm long, to 1 mm wide. Fronds crowded, erect, caespitose, homomorphic, to 160 mm long; stipe firm, proximally brown, green higher up, narrowly winged along entire length, shallowly sulcate adaxially, to 130 mm long, to 1.2 mm in diameter, sparsely scaled, scales thin-chartaceus, concolorous, stramineous, sessile, linear-hastate, entire or irregularly set with short marginal outgrowths, apex terminates in an oblong thin-walled cell, to 3 mm long, to 0.5 mm wide; lamina anadromous, firmly herbaceous, flabellate, edges forming an angle of 150-180°, divided into 2 symmetrical halves by a deep median notch, each to 4 times dichotomously branched, to 32 mm long, muricate adaxially, abaxially initially sparsely set with scales similar to, but smaller than, those on the stipe, lamina declinate when desiccated; segments up to 48 per lamina, linear, of variable length, to 1.2 mm wide, each ending in 3 or 4 acute teeth. Venation adaxially obscure, raised abaxially, dichotomously forked, ending in the teeth. Sori linear, along outer segment veins; receptacle nude; sporangium stalk simple, 3-seriate below capsule, capsule broadly elliptic in lateral view, with (17-)19(-20) indurated annulus cells, epistomium (5-)6(-7)-celled, hypostomium (5-)6(-6)-celled; indusium membranous, continuous along segment margin, entire. Spores trilete, tetrahedral-globose, with an equatorial flange, proximal face verrucuose, distal face hemispherical, rugate, (46-)56.2(-60) (m in diameter. Figure 15C.