Plants terrestrial or epilithic. Rhizome erect to suberect, to 20 mm long, to 4 mm in diameter, set with roots, crowded stipe bases and scales, scales chartaceus, atrocastaneus, clathrate, sessile, subulate to narrowly lanceolate, cordate, margins irregularly set with short filiform outgrowths, apex terminates in an oblong thin-walled cell, to 5 mm long, to 0.7 mm wide. Fronds crowded, caespitose, erect to arching, to 475 mm long; stipe firm, castaneus, terete, to 120 mm long, to 1.2 mm in diameter, narrowly green-winged for most of the length, initially moderately set with hairs and scales, subglabrous later, hairs 3-5-celled, clavate, to 0.25 mm long, scales chartaceus, atrocastaneus, clathrate, stalked, subulate to filiform, the larger often with short filiform outgrowths along the margin, apex terminates in an oblong thin-walled cell, to 5 mm long, to 0.7 mm wide; lamina anadromous, 2-pinnate, narrowly elliptic, to 360 mm long, to 85 mm wide, with up to 23 petiolated pinna pairs; rachis firm, castaneus, green towards the apex, terete, narrowly green-winged, sparsely set with hairs and scales similar to those on the stipe, glabrous later; pinnae petiolate, petiole to 1 mm long, opposite to alternate, basally more widely spaced than apically, pinnae in the lower half gradually reduced, ovate to narrowly ovate, to 45 mm long, to 16 mm wide, with up to 5 pinnule pairs; pinna-rachis terete, narrowly winged, glabrous; pinnules herbaceous, sessile, rhombic to flabellate, coarsely serrate or divided into 2-fid or 3-fid lobes, lobe apices entire or shallowly repand to erose, to 12 mm long, to 10 mm wide, glabrous adaxially and abaxially. Venation anadromous, evident, flabellately forked, ending in the teeth near the margin. Sori linear, usually on the terminal vein branches, but often also at a fork, to 3.5 mm long; indusium chartaceus, stramineous, linear, entire, to 3.5 mm long, to 0.3 mm wide; sporangium long-stalked, uniseriate, 3-seriate below the capsule, capsule globose to broadly elliptic in lateral view, annulus with (17-)19(-19) indurated annulus cells, epistomium 2-celled, hypostomium 4-celled. Spores 64 per sporangium, brown, elliptic, monolete, perispore forms broad reticulate wings, exospore (28-)31.3(-34) x (18-)20.1(-22) (m. Figure 48E & F.