Information

Plants terrestrial or on rock. Stems creeping, ascending, or erect, stolons absent. Leaves monomorphic, evergreen or dying back in winter. Petiole ca. 1/2 to equaling length of blade, base swollen and persisting as trophopod over winter or not; vascular bundles 2, lateral, lunate in cross section. Blade oblong-lanceolate to deltate, 1-pinnate to 2-pinnate-pinnatifid [simple to 4-pinnate-pinnatifid], gradually reduced distally to pinnatifid apex or apical pinna similar to (conform) adjacent pinnae, herbaceous to papery. Pinnae not articulate to rachis, segment margins entire, crenulate, or serrate; proximal pinnae not reduced, sessile, equilateral or inequilateral; costae adaxially deeply grooved, grooves continuous with that of rachis; indument abaxially absent or of linear to ovate scales, adaxially absent. Veins free, simple or forked, or basal pairs of adjacent segments anastomosing. Sori single or paired back-to-back on veins, oblong to linear, straight or slightly falcate; indusia linear, laterally attached, persistent. Spores brownish, usually broadly winged. x = 40, 41.