Subshrubs, perennial, gray-green, grayish, or sometimes reddish, to 60 cm, canescent, pruinose, or densely lanuginose to glabrate; trichomes completely or partially candelabriform. Stems ascending or decumbent, suffrutescent or suffruticulose, buds commonly present, rarely absent, on stem bases, lanate or densely lanate. Leaves: petiole to 2.5 cm; blade lanceolate, ovate, ovate-oblong, very widely ovate, circular, or reniform, 0.6–4.5 × 0.4–2.7 cm, chartaceous, base cordate, cuneate, lightly attenuate, or oblique. Inflorescences 1–3(–4)-flowered; involucral leaf petiole to 0.9 cm, blade lanceolate, ovate, very widely ovate, circular, or reniform, 0.2–2.7 × 0.2–2.1 cm, chartaceous, base cuneate, cordate, attenuate, or oblique, apex acute or obtuse; involucre on secondary branches formed by connation of involucral leaf petioles, which becomes indurate and adnate with stem or with leaf petiole and stem; bracts widely ovate or widely depressed-ovate, 0.8–1.6 × 0.8–1.6 mm, apex acute or obtuse, crenate, lanuginose distally or glabrous; bracteoles 0.9–1.5 × 0.6–1.4 mm, apex acute or obtuse, lanuginose or glabrous. Flowers 1.7–3 mm; tepals yellowish or yellowish brown, 1.7–2.7 × 0.6–1.3 mm, densely lanuginose to glabrous; staminal cup 0.4–0.9 mm; filaments 0.5–1 mm; anthers 0.5–0.9 mm; pseudostaminodes absent or triangular lobes, 0.1–0.3 mm; ovary 0.3–0.7 × 0.5–0.9 mm; style to 0.1 mm; stigmas 0.2–0.5 mm. Utricles 1–1.8 × 0.8–1.6 mm. Seeds brown-red, 0.9–1.5 × 0.8–1.4 mm.