Plants 0.2–1.5 m. Stems erect; branches brownish or red-brown, (dull), glabrous, long-silky, or villous to glabrescent; branchlets yellow-brown or gray-brown, long-silky, pilose, or moderately densely villous. Leaves: stipules absent or rudimentary; petiole (shallowly to deeply grooved adaxially), 2–8 mm, (with 2 spherical glands distally, dark brown, sometimes basilaminar, sparsely pubescent or glabrous adaxially); largest medial blade hypostomatous, (veins strongly impressed-reticulate), broadly elliptic, subcircular, or obovate, 18–67 × 10–40 mm, 1.1–2.3 times as long as wide, base rounded, convex, or subcordate, margins strongly revolute, crenate or subentire, apex rounded, convex, retuse, or toothed, abaxial surface sparsely to densely villous or long-silky, veins often with long, straight hairs, adaxial slightly glossy, glabrous or sparsely long-silky; proximal blade margins entire or crenate; juvenile blade (yellowish green), abaxially very densely long-silky. Catkins: staminate 13–48 × 4–6.5(–8) mm, flowering branchlet 3–31(–50) mm; pistillate densely flowered, slender or stout, 18–56 × 4–10 mm, flowering branchlet 3–27(–40) mm; floral bract tawny, 0.8–1.6 mm, apex rounded, entire, abaxially densely hairy, hairs straight. Staminate flowers: abaxial nectary 0.6–0.8 mm, adaxial nectary narrowly oblong, 0.5–1.2 mm, nectaries connate and shallowly cup-shaped, or distinct; filaments distinct, hairy on proximal 1/2; anthers ellipsoid or globose, 0.3–0.5 mm. Pistillate flowers: abaxial nectary (0–)0.8–0.9 mm, adaxial nectary oblong, ovate, or narrowly oblong to almost filiform, 0.7–1.4 mm, shorter to longer than stipe, nectaries distinct or connate and cup-shaped; stipe 0.4–1.2 mm; ovary pyriform or obnapiform, densely short-silky, hairs cylindrical, beak abruptly tapering to styles; ovules 13–15 per ovary; styles connate 1/2 their lengths to almost distinct, 0.2–0.4 mm; stigmas flat, abaxially non-papillate with rounded tip, slenderly cylindrical, or 2 plump lobes, 0.2–0.28–0.36 mm. Capsules 3–5 mm. 2n = 38.
