Perennials; puberulent, (trichomes cylindrical). Stems simple from base, erect, branched (several) distally, (0.6–)1–5.3(–7.7) dm. Basal leaves (soon withered); not rosulate; petiole 1–4 cm; blade oblanceolate or pinnatifid, 2–5 cm × 8–15 mm, margins (of lobes) entire serrate or dentate. Cauline leaves shortly petiolate or sessile; blade oblanceolate or (distal) linear, (0.6–)1.2–4.8(–6) cm × 1–8(–10) mm, base attenuate to cuneate, not auriculate, margins dentate, (distal) entire, or, rarely, lobed. Racemes slightly elongated in fruit; rachis puberulent, trichomes curved, cylindrical to subclavate. Fruiting pedicels divaricate-ascending to horizontal, straight or recurved, (terete), (1.6–)2–3.8(–5) × 0.2–0.3 mm, usually puberulent adaxially, rarely throughout. Flowers: sepals oblong, 0.6–0.9(–1.1) × 0.3–0.4 mm; petals (absent or rudimentary), white, linear, 0.2–0.8(–1) × 0.05–0.1 mm, claw absent; stamens 2, median; filaments 0.6–0.9 mm; anthers 0.15–0.2 mm. Fruits elliptic, 2.2–3.2 × 1.7–2.1mm, apically winged, apical notch 0.1–0.3(–0.4) mm deep; valves thin, smooth, not veined, glabrous or puberulent at least along margin; style usually obsolete, rarely to 0.1 mm, included in apical notch. Seeds oblong, 1.2–1.6 × 0.8–0.9 mm. 2n = 32, 64.