Leaves ovate to oblong, apex broadly acute or occasionally rounded, apiculate or short- to long-mucronate, occasionally short-awned, margins recurved in middle 2/3 of leaf or occasionally almost plane, not or very weakly bordered distally with ca. 2 rows of rhomboid, less papillose cells; costa subpercurrent, percurrent or excurrent, lacking an adaxial pad of cells, distally narrow, 2–3 cells across the convex adaxial surface; distal laminal cells hexagonal, 15–20 µm wide, 1:1, densely papillose with several 2-fid papillae. Sexual condition autoicous. Sporophytes exerted. Seta 0.9–1.2(–1.9) cm. Capsule stegocarpic, rarely systylius, cylindric, erect and nearly straight, urn 1.5–2(–2.8) mm; peristome (150–)300–350 length µm, teeth 16, usually not twisted, divided to near the base or perforate basally, the teeth often basally fused in 2s or 4s, basal membrane very low; operculum 0.7–1 mm. Spores ca. 20–23(–25) µm, spheric or elliptic, densely papillose.
