Shrubs or trees, 0.5–5(–10) m. Stems: branches yellow-brown to red-brown; branchlets yellowish, yellow-brown, or red-brown, puberulent, densely short-silky-tomentose or short-silky-villous, hairs appressed, sometimes few spreading. Leaves: stipules absent or rudimentary on early ones, foliaceous on late ones; petiole pubescent adaxially; largest medial blade lorate, 30–136–143 × 2–8 mm, 10–16–28 times as long as wide, abaxial surface densely silky-villous or pilose, hairs appressed or spreading, straight or wavy, adaxial sparsely long-silky to glabrescent; juvenile blade very densely long-silky abaxially (hairs usually appressed, sometimes a few spreading). Catkins: staminate 13–54 × 2–10 mm, flowering branchlet 1.5–16 mm; pistillate loosely flowered, slender or stout, 14.5–70 × 4–12 mm, flowering branchlet 2–16 mm; floral bract 1.2–1.6 mm, apex rounded, entire, abaxially hairy (or glabrate), hairs wavy or straight. Staminate flowers: abaxial nectary 0.2–0.8 mm, adaxial nectary oblong, 0.3–0.8 mm; anthers 0.5–1 mm. Pistillate flowers: adaxial nectary oblong or ovate, 0.3–0.9 mm shorter than stipe; stipe 0.2–0.9 mm; ovary obclavate, usually glabrous, sometimes beak pilose, beak bulged below or abruptly tapering to styles; ovules 13–30 per ovary; styles 0–0.2 mm; stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.3–0.5 mm. 2n = 38.