Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pub-escent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). Stems several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. Basal leaves: (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). Cauline leaves: blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). Racemes loose, (elongated in fruit). Fruiting pedicels (divaricate, slightly sigmoid or nearly straight), 6–12 mm. Flowers: sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. Fruits (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. Seeds (dark brown), flattened, (2–3 mm). 2n = 10, 16, 24.
